Philip Meneely (PhD, U of Minnesota) is a Professor of Biology at Haverford College where he has taught both introductory and advanced genetics for more than 20 years, as well as courses in genomics and bioinformatics. He previously was on the faculty of the Fred Hutchinson Cancer Research Center. His research with C. elegans has included publications on chromosome rearrangements, polyploidy, meiosis, sex determination, dosage compensation, and gene interactions. He is also the author of Genetic Analysis: Genes, Genomes, and Networks in Eukaryotes (Oxford University Press), now in its second edition, which was short-listed by the Royal Society (London) in 2015 for Undergraduate Biology Textbook of the year.

Rachel Dawes Hoang (Ph.D. Cambridge University, UK) is an Associate Professor in the Biology Department at Haverford College. She has published research and review articles in the fields of developmental biology and evolutionary developmental biology. Her current research investigates the evolution of genes controlling cell shape changes as well as the interactions between endosymbiotic bacteria and host cells during embryonic development of insects. She regularly teaches courses in genetics, evolution, and development. She is currently the chair of the Biology Department. She is a former Helen Hay Whitney fellow.

Iruka N. Okeke (PhD, Obafemi Awolowo University, Nigeria) taught biology at Haverford College, PA, USA from 2002 until 2014. She is presently Professor of Pharmaceutical Microbiology at the University of Ibadan, Nigeria and has also taught in other African and UK applied health programs. Her research on bacterial genetics and microbiology focuses on intestinal pathogens and on antimicrobial resistance. She is co-author of two books and about a hundred articles and chapters. She has been the recipient of Fulbright, Branco Weiss and Institute for Advanced Study (Berlin) Fellowships. Okeke serves on editorial, higher education, health policy and science policy advisory panels and boards in the US, Europe and Africa.

Katherine Heston (M.S. University of Wisconsin, A.B. Princeton University) is an Instructor in Biology at Haverford College. She has been involved in teaching undergraduates for thirty years at Lake Forest College (IL), Northwestern University, Villanova University, and Haverford College. Her teaching background includes botany, ecology, genetics, cell and molecular biology. Working closely with students in the teaching lab has developed her sense of the student perspective, which informs her development of effective teaching materials for her classes.

• 1: Darwin's Finches: Evolution, Genomes, and Genes


• 2: The Central Dogma of Molecular Biology


• 3: Genome Structure, Organization, and Variation


• 4: Descent with Modification: DNA Replication and Mutation


• 5: The Inheritance of Single Gene Traits


• 6: The Cellular Basis for Mendelian Genetics


• 7: X-linked Genes and Sex Chromosomes


• 8: The Inheritance of Multiple Genes


• 9: The Locations of Genes on Chromosomes: Linkage and Genetic Maps


• 10: Human Genetic Mapping, Genome Wide Association Studies, and Complex Traits


• 11: Exchange and Evolution


• 12: Transcription: Reading and Expressing Genes


• 13: Translation: From Nucleic Acids to Amino Acids


• 14: Networks of Gene Regulation


• 15: Genetic Analysis of Cellular Processes


• 16: The Genetics of Populations


• 17: Metagenomes: Genome Analysis of Communities